# Branch Survey — 10 October 2026

Screened 1,308 policy-selected candidates against the full PBDB classification captured on 9 October. This is not a list of approved viable answers.

108 direct-parent groups have at least five genera. Matching diet, period and region evidence produces 63 provisional pools across 53 parents, covering 484 genera. 60 pools could reach five remaining candidates; three isolated five-member pools fall to four after their first within-group guess.

## All matching pools

| Direct parent | Source diet | Periods | Regions | Before guesses | After one member guessed |
| --- | --- | --- | --- | --- | --- |
| Hadrosauroidea | herbivore | Cretaceous | Asia | 24 | 23 |
| Nodosauridae | herbivore | Cretaceous | Americas | 20 | 19 |
| Mamenchisauridae | herbivore | Jurassic | Asia | 19 | 18 |
| Chasmosaurinae | herbivore | Cretaceous | Americas | 16 | 15 |
| Euhelopodidae | herbivore | Cretaceous | Asia | 15 | 14 |
| Titanosauria | herbivore | Cretaceous | Americas | 12 | 11 |
| Heyuanninae | omnivore | Cretaceous | Asia | 11 | 10 |
| Parvicursorinae | carnivore | Cretaceous | Asia | 11 | 10 |
| Rebbachisauridae | herbivore | Cretaceous | Americas | 11 | 10 |
| Rinconsauria | herbivore | Cretaceous | Americas | 11 | 10 |
| Elasmaria | herbivore | Cretaceous | Americas | 10 | 9 |
| Ankylosaurinae | herbivore | Cretaceous | Asia | 9 | 8 |
| Neoceratopsia | herbivore | Cretaceous | Asia | 9 | 8 |
| Somphospondyli | herbivore | Cretaceous | Asia | 9 | 8 |
| Therizinosauridae | herbivore | Cretaceous | Asia | 9 | 8 |
| Troodontidae | carnivore, omnivore | Cretaceous | Asia | 9 | 8 |
| Elasmaria | herbivore | Cretaceous | Oceania | 8 | 7 |
| Leptoceratopsidae | herbivore | Cretaceous | Americas | 8 | 7 |
| Lithostrotia | herbivore | Cretaceous | Americas | 8 | 7 |
| Somphospondyli | herbivore | Cretaceous | Americas | 8 | 7 |
| Ankylosauridae | herbivore | Cretaceous | Asia | 7 | 6 |
| Centrosaurinae | herbivore | Cretaceous | Americas | 7 | 6 |
| Furileusauria | carnivore | Cretaceous | Americas | 7 | 6 |
| Hadrosauroidea | herbivore | Cretaceous | Americas | 7 | 6 |
| Kritosaurini | herbivore | Cretaceous | Americas | 7 | 6 |
| Megaraptoridae | carnivore | Cretaceous | Americas | 7 | 6 |
| Microraptorinae | carnivore | Cretaceous | Asia | 7 | 6 |
| Sauropoda | herbivore | Jurassic | Asia | 7 | 6 |
| Titanosauria | herbivore | Cretaceous | Asia | 7 | 6 |
| Aeolosaurini | herbivore | Cretaceous | Americas | 6 | 5 |
| Brachyrostra | carnivore | Cretaceous | Americas | 6 | 5 |
| Caenagnathinae | omnivore | Cretaceous | Americas | 6 | 5 |
| Carcharodontosauridae | carnivore | Cretaceous | Americas | 6 | 5 |
| Citipatinae | omnivore | Cretaceous | Asia | 6 | 5 |
| Dromaeosauridae | carnivore | Cretaceous | Asia | 6 | 5 |
| Eusauropoda | herbivore | Jurassic | Asia | 6 | 5 |
| Eutitanosauria | herbivore | Cretaceous | Americas | 6 | 5 |
| Lognkosauria | herbivore | Cretaceous | Americas | 6 | 5 |
| Ornithomimidae | herbivore, omnivore | Cretaceous | Americas | 6 | 5 |
| Ornithomimidae | herbivore, omnivore | Cretaceous | Asia | 6 | 5 |
| Tyrannosauroidea | carnivore | Cretaceous | Americas | 6 | 5 |
| Tyrannosauroidea | carnivore | Cretaceous | Asia | 6 | 5 |
| Unenlagiinae | carnivore | Cretaceous | Americas | 6 | 5 |
| Abelisauridae | carnivore | Cretaceous | Americas | 5 | 4 |
| Alvarezsauridae | carnivore | Cretaceous | Americas | 5 | 4 |
| Alvarezsauroidea | carnivore | Cretaceous | Asia | 5 | 4 |
| Arenysaurini | herbivore | Cretaceous | Europe | 5 | 4 |
| Brachylophosaurini | herbivore | Cretaceous | Americas | 5 | 4 |
| Centrosaurini | herbivore | Cretaceous | Americas | 5 | 4 |
| Dicraeosauridae | herbivore | Jurassic | Americas | 5 | 4 |
| Dromaeosauridae | carnivore | Cretaceous | Europe | 5 | 4 |
| Eutitanosauria | herbivore | Cretaceous | Europe | 5 | 4 |
| Leptoceratopsidae | herbivore | Cretaceous | Asia | 5 | 4 |
| Nasutoceratopsini | herbivore | Cretaceous | Americas | 5 | 4 |
| Neornithischia | herbivore | Jurassic | Asia | 5 | 4 |
| Nodosauridae | herbivore | Cretaceous | Europe | 5 | 4 |
| Orodrominae | herbivore | Cretaceous | Americas | 5 | 4 |
| Oviraptorosauria | omnivore | Cretaceous | Asia | 5 | 4 |
| Pachycephalosaurinae | herbivore | Cretaceous | Americas | 5 | 4 |
| Paraves | carnivore | Jurassic | Asia | 5 | 4 |
| Saltasaurinae | herbivore | Cretaceous | Americas | 5 | 4 |
| Styracosterna | herbivore | Cretaceous | Europe | 5 | 4 |
| Therizinosauria | carnivore | Cretaceous | Asia | 5 | 4 |

## Genus membership

### Hadrosauroidea · herbivore · Cretaceous · Asia · 24 candidates

Datonglong, Equijubus, Fukuisaurus, Gilmoreosaurus, Gobihadros, Gongpoquansaurus, Haolong, Jintasaurus, Koshisaurus, Levnesovia, Nanningosaurus, Nanyangosaurus, Penelopognathus, Plesiohadros, Probactrosaurus, Qianjiangsaurus, Ratchasimasaurus, Shuangmiaosaurus, Sirindhorna, Tanius, Xuwulong, Yunganglong, Zhanghenglong, Zuoyunlong.

### Nodosauridae · herbivore · Cretaceous · Americas · 20 candidates

Acantholipan, Animantarx, Borealopelta, Edmontonia, Gastonia, Glyptodontopelta, Hoplitosaurus, Invictarx, Niobrarasaurus, Nodosaurus, Panoplosaurus, Patagopelta, Pawpawsaurus, Peloroplites, Propanoplosaurus, Sauropelta, Silvisaurus, Stegopelta, Tatankacephalus, Texasetes.

### Mamenchisauridae · herbivore · Jurassic · Asia · 19 candidates

Abrosaurus, Analong, Anhuilong, Bellusaurus, Chuanjiesaurus, Daanosaurus, Eomamenchisaurus, Huangshanlong, Hudiesaurus, Jingiella, Klamelisaurus, Omeisaurus, Qijianglong, Rhomaleopakhus, Tienshanosaurus, Tonganosaurus, Tongnanlong, Xinjiangtitan, Yuanmousaurus.

### Chasmosaurinae · herbivore · Cretaceous · Americas · 16 candidates

Agujaceratops, Anchiceratops, Arrhinoceratops, Bisticeratops, Bravoceratops, Chasmosaurus, Coahuilaceratops, Judiceratops, Kosmoceratops, Navajoceratops, Pentaceratops, Sierraceratops, Spiclypeus, Terminocavus, Utahceratops, Vagaceratops.

### Euhelopodidae · herbivore · Cretaceous · Asia · 15 candidates

Daxiatitan, Erketu, Euhelopus, Gobititan, Huabeisaurus, Nagatitan, Phuwiangosaurus, Qiaowanlong, Ruixinia, Ruyangosaurus, Silutitan, Tambatitanis, Tangvayosaurus, Xianshanosaurus, Yongjinglong.

### Titanosauria · herbivore · Cretaceous · Americas · 12 candidates

Andesaurus, Argyrosaurus, Baalsaurus, Barrosasaurus, Choconsaurus, Laplatasaurus, Malarguesaurus, Narambuenatitan, Ninjatitan, Petrobrasaurus, Tiamat, Yeneen.

### Heyuanninae · omnivore · Cretaceous · Asia · 11 candidates

Banji, Conchoraptor, Ganzhousaurus, Heyuannia, Ingenia, Jiangxisaurus, Khaan, Machairasaurus, Nemegtomaia, Oksoko, Shixinggia.

### Parvicursorinae · carnivore · Cretaceous · Asia · 11 candidates

Albinykus, Ceratonykus, Dzharaonyx, Jaculinykus, Khulsanurus, Linhenykus, Mononykus, Ondogurvel, Parvicursor, Shuvuuia, Xixianykus.

### Rebbachisauridae · herbivore · Cretaceous · Americas · 11 candidates

Agustinia, Amazonsaurus, Astigmasaura, Campananeyen, Cienciargentina, Comahuesaurus, Katepensaurus, Lavocatisaurus, Rayososaurus, Sidersaura, Zapalasaurus.

### Rinconsauria · herbivore · Cretaceous · Americas · 11 candidates

Arrudatitan, Brasilotitan, Bravasaurus, Chadititan, Maxakalisaurus, Muyelensaurus, Overosaurus, Pitekunsaurus, Punatitan, Rinconsaurus, Uberabatitan.

### Elasmaria · herbivore · Cretaceous · Americas · 10 candidates

Anabisetia, Chakisaurus, Gasparinisaura, Isasicursor, Macrogryphosaurus, Mahuidacursor, Notohypsilophodon, Sektensaurus, Talenkauen, Tietasaura.

### Ankylosaurinae · herbivore · Cretaceous · Asia · 9 candidates

Crichtonpelta, Eopinacosaurus, Jinyunpelta, Pinacosaurus, Saichania, Talarurus, Tarchia, Tsagantegia, Zaraapelta.

### Neoceratopsia · herbivore · Cretaceous · Asia · 9 candidates

Archaeoceratops, Asiaceratops, Auroraceratops, Beg, Graciliceratops, Liaoceratops, Mosaiceratops, Sasayamagnomus, Yamaceratops.

### Somphospondyli · herbivore · Cretaceous · Asia · 9 candidates

Dongbeititan, Dzharatitanis, Gannansaurus, Huanghetitan, Jiangshanosaurus, Jiangxititan, Liubangosaurus, Sonidosaurus, Yunmenglong.

### Therizinosauridae · herbivore · Cretaceous · Asia · 9 candidates

Duonychus, Enigmosaurus, Erliansaurus, Nanshiungosaurus, Paralitherizinosaurus, Segnosaurus, Suzhousaurus, Therizinosaurus, Tiantaisaurus.

### Troodontidae · carnivore, omnivore · Cretaceous · Asia · 9 candidates

Almas, Archaeornithoides, Daliansaurus, Gobivenator, Harenadraco, Papiliovenator, Sinusonasus, Tochisaurus, Xixiasaurus.

### Elasmaria · herbivore · Cretaceous · Oceania · 8 candidates

Atlascopcosaurus, Diluvicursor, Fostoria, Galleonosaurus, Leaellynasaura, Muttaburrasaurus, Qantassaurus, Weewarrasaurus.

### Leptoceratopsidae · herbivore · Cretaceous · Americas · 8 candidates

Cerasinops, Ferrisaurus, Gremlin, Gryphoceratops, Leptoceratops, Montanoceratops, Prenoceratops, Unescoceratops.

### Lithostrotia · herbivore · Cretaceous · Americas · 8 candidates

Arackar, Atacamatitan, Bustingorrytitan, Elaltitan, Epachthosaurus, Kaijutitan, Nullotitan, Pellegrinisaurus.

### Somphospondyli · herbivore · Cretaceous · Americas · 8 candidates

Astrophocaudia, Brontomerus, Chubutisaurus, Dasosaurus, Ligabuesaurus, Padillasaurus, Sauroposeidon, Triunfosaurus.

### Ankylosauridae · herbivore · Cretaceous · Asia · 7 candidates

Bissektipelta, Chuanqilong, Datai, Gobisaurus, Huaxiazhoulong, Liaoningosaurus, Shamosaurus.

### Centrosaurinae · herbivore · Cretaceous · Americas · 7 candidates

Diabloceratops, Machairoceratops, Menefeeceratops, Monoclonius, Stellasaurus, Wendiceratops, Xenoceratops.

### Furileusauria · carnivore · Cretaceous · Americas · 7 candidates

Carnotaurus, Elemgasem, Koleken, Llukalkan, Pycnonemosaurus, Quilmesaurus, Viavenator.

### Hadrosauroidea · herbivore · Cretaceous · Americas · 7 candidates

Cedrorestes, Claosaurus, Eolambia, Gonkoken, Huehuecanauhtlus, Jeyawati, Protohadros.

### Kritosaurini · herbivore · Cretaceous · Americas · 7 candidates

Coahuilasaurus, Gryposaurus, Huallasaurus, Kelumapusaura, Kritosaurus, Naashoibitosaurus, Secernosaurus.

### Megaraptoridae · carnivore · Cretaceous · Americas · 7 candidates

Aerosteon, Joaquinraptor, Maip, Megaraptor, Murusraptor, Orkoraptor, Tratayenia.

### Microraptorinae · carnivore · Cretaceous · Asia · 7 candidates

Changyuraptor, Graciliraptor, Jian, Microraptor, Shanag, Sinornithosaurus, Zhongjianosaurus.

### Sauropoda · herbivore · Jurassic · Asia · 7 candidates

Chinshakiangosaurus, Damalasaurus, Isanosaurus, Kotasaurus, Lamplughsaura, Microdontosaurus, Sanpasaurus.

### Titanosauria · herbivore · Cretaceous · Asia · 7 candidates

Abdarainurus, Baotianmansaurus, Borealosaurus, Dongyangosaurus, Gandititan, Hamititan, Mongolosaurus.

### Aeolosaurini · herbivore · Cretaceous · Americas · 6 candidates

Aeolosaurus, Baurutitan, Caieiria, Gondwanatitan, Mesetasaurus, Panamericansaurus.

### Brachyrostra · carnivore · Cretaceous · Americas · 6 candidates

Aucasaurus, Ekrixinatosaurus, Guemesia, Ilokelesia, Skorpiovenator, Thanos.

### Caenagnathinae · omnivore · Cretaceous · Americas · 6 candidates

Anzu, Apatoraptor, Caenagnathus, Citipes, Epichirostenotes, Hagryphus.

### Carcharodontosauridae · carnivore · Cretaceous · Americas · 6 candidates

Acrocanthosaurus, Giganotosaurus, Lajasvenator, Mapusaurus, Taurovenator, Unquillosaurus.

### Citipatinae · omnivore · Cretaceous · Asia · 6 candidates

Citipati, Corythoraptor, Huanansaurus, Rinchenia, Tongtianlong, Wulatelong.

### Dromaeosauridae · carnivore · Cretaceous · Asia · 6 candidates

Daurlong, Kuru, Luanchuanraptor, Tianyuraptor, Wulong, Zhenyuanlong.

### Eusauropoda · herbivore · Jurassic · Asia · 6 candidates

Datousaurus, Ferganasaurus, Huashanosaurus, Jinchuanloong, Nebulasaurus, Shunosaurus.

### Eutitanosauria · herbivore · Cretaceous · Americas · 6 candidates

Argentinosaurus, Bonitasaura, Dreadnoughtus, Inawentu, Menucocelsior, Patagotitan.

### Lognkosauria · herbivore · Cretaceous · Americas · 6 candidates

Austroposeidon, Drusilasaura, Futalognkosaurus, Mendozasaurus, Quetecsaurus, Traukutitan.

### Ornithomimidae · herbivore, omnivore · Cretaceous · Americas · 6 candidates

Dromiceiomimus, Mexidracon, Ornithomimus, Rativates, Struthiomimus, Tototlmimus.

### Ornithomimidae · herbivore, omnivore · Cretaceous · Asia · 6 candidates

Aepyornithomimus, Anserimimus, Dzharacursor, Gallimimus, Qiupalong, Sinornithomimus.

### Tyrannosauroidea · carnivore · Cretaceous · Americas · 6 candidates

Appalachiosaurus, Dryptosaurus, Gualicho, Moros, Suskityrannus, Teihivenator.

### Tyrannosauroidea · carnivore · Cretaceous · Asia · 6 candidates

Dilong, Jinbeisaurus, Khankhuuluu, Raptorex, Timurlengia, Xiongguanlong.

### Unenlagiinae · carnivore · Cretaceous · Americas · 6 candidates

Austroraptor, Buitreraptor, Diuqin, Neuquenraptor, Unenlagia, Ypupiara.

### Abelisauridae · carnivore · Cretaceous · Americas · 5 candidates

Kurupi, Spectrovenator, Tralkasaurus, Vitosaura, Xenotarsosaurus.

### Alvarezsauridae · carnivore · Cretaceous · Americas · 5 candidates

Achillesaurus, Albertonykus, Alnashetri, Alvarezsaurus, Trierarchuncus.

### Alvarezsauroidea · carnivore · Cretaceous · Asia · 5 candidates

Bannykus, Nemegtonykus, Qiupanykus, Tugulusaurus, Xiyunykus.

### Arenysaurini · herbivore · Cretaceous · Europe · 5 candidates

Adynomosaurus, Arenysaurus, Blasisaurus, Canardia, Pararhabdodon.

### Brachylophosaurini · herbivore · Cretaceous · Americas · 5 candidates

Acristavus, Brachylophosaurus, Maiasaura, Ornatops, Probrachylophosaurus.

### Centrosaurini · herbivore · Cretaceous · Americas · 5 candidates

Centrosaurus, Coronosaurus, Rubeosaurus, Spinops, Styracosaurus.

### Dicraeosauridae · herbivore · Jurassic · Americas · 5 candidates

Athenar, Dyslocosaurus, Kaatedocus, Smitanosaurus, Suuwassea.

### Dromaeosauridae · carnivore · Cretaceous · Europe · 5 candidates

Balaur, Dromaeosauroides, Nuthetes, Pyroraptor, Variraptor.

### Eutitanosauria · herbivore · Cretaceous · Europe · 5 candidates

Lohuecotitan, Magyarosaurus, Normanniasaurus, Paludititan, Petrustitan.

### Leptoceratopsidae · herbivore · Cretaceous · Asia · 5 candidates

Helioceratops, Ischioceratops, Koreaceratops, Udanoceratops, Zhuchengceratops.

### Nasutoceratopsini · herbivore · Cretaceous · Americas · 5 candidates

Avaceratops, Crittendenceratops, Furcatoceratops, Nasutoceratops, Yehuecauhceratops.

### Neornithischia · herbivore · Jurassic · Asia · 5 candidates

Agilisaurus, Hexinlusaurus, Pulaosaurus, Sanxiasaurus, Yandusaurus.

### Nodosauridae · herbivore · Cretaceous · Europe · 5 candidates

Anoplosaurus, Hungarosaurus, Hylaeosaurus, Polacanthus, Struthiosaurus.

### Orodrominae · herbivore · Cretaceous · Americas · 5 candidates

Albertadromeus, Nevadadromeus, Orodromeus, Oryctodromeus, Zephyrosaurus.

### Oviraptorosauria · omnivore · Cretaceous · Asia · 5 candidates

Caudipteryx, Incisivosaurus, Ningyuansaurus, Protarchaeopteryx, Yuanyanglong.

### Pachycephalosaurinae · herbivore · Cretaceous · Americas · 5 candidates

Acrotholus, Brontotholus, Foraminacephale, Platytholus, Sphaerotholus.

### Paraves · carnivore · Jurassic · Asia · 5 candidates

Caihong, Eosinopteryx, Epidexipteryx, Pedopenna, Serikornis.

### Saltasaurinae · herbivore · Cretaceous · Americas · 5 candidates

Alamosaurus, Bonatitan, Ibirania, Rocasaurus, Yamanasaurus.

### Styracosterna · herbivore · Cretaceous · Europe · 5 candidates

Calvarius, Delapparentia, Istiorachis, Magnamanus, Riabininohadros.

### Therizinosauria · carnivore · Cretaceous · Asia · 5 candidates

Erlikosaurus, Fukuivenator, Jianchangosaurus, Kayrasaurus, Lingyuanosaurus.

## Limitations and checks

- These are candidate pools, not confirmed viable hidden answers. Diet and occurrence evidence needs review.
- Most source diets are inherited from higher taxa. Mixed categories are preserved, not forced into one gameplay category.
- Occurrence periods are the union of positive overlaps with PBDB period bounds, not a claim of continuous genus survival.
- Broad regions come from modern country codes; RU, KZ and offshore O2 remain unresolved. Missing information never acts as a matching clue.
- All linked occurrences form the main screen; excluding cf., aff. and ? records is a sensitivity check, not an approved occurrence policy.
- Counts use the full PBDB classification. Omitting source groups can produce larger crowds.
- Pool sizes precede guesses; the automatic rule counts five unguessed possibilities after every observed relationship and requested clue.
- Only the existing 36-genus sample is playable; this audit does not expand the live roster.

- 131 genera have unresolved screening fields: 76 missing diets, 24 missing occurrences and 33 locality-mapping reviews. Counts overlap.
- The explicitly unqualified-occurrence sensitivity screen gives 65 pools and 496 genera; the five largest pool memberships are unchanged.
- Using only current prototype clade names as a diagnostic source-tree projection yields 71 pools and 836 genera. Preserve useful PBDB branches when expanding.

## Next review

The Locality Survey now covers all 108 large parent groups and all 63 matching pools. See LOCALITY-SURVEY.md and locality-survey.html for occurrence overlaps, candidate splits and evidence needing review. No live roster or answer eligibility is changed.